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L'Abruzz ruggisce!
Published: 2017/02/11
Channel: RedZone
Lancian family
Lancian family
Published: 2014/10/02
Channel: Jhonrick Lancian
Lancian family
Lancian family
Published: 2014/10/02
Channel: Jhonrick Lancian
dennis morales, leyes boy lancian, pearl john lancian bodutz warriors
dennis morales, leyes boy lancian, pearl john lancian bodutz warriors
Published: 2014/10/25
Channel: elner ray aloy
my family Lancian(1)
my family Lancian(1)
Published: 2015/04/05
Channel: saihane lancian
Lancian Galaxy
Lancian Galaxy
Published: 2015/10/24
Channel: Warner/Chappell Productions - Topic
Marco Papa - TROY in pescarese: LANCIANO in serie B con il PESCARA (Iachille)
Marco Papa - TROY in pescarese: LANCIANO in serie B con il PESCARA (Iachille)
Published: 2015/02/06
Channel: Lorenzo Cav95
14 Alla feria di Lanciana
14 Alla feria di Lanciana
Published: 2013/07/13
Channel: David Neaud
I LOVE YOU(2NE1)
I LOVE YOU(2NE1)
Published: 2015/01/25
Channel: christianjames lancian
LANCIAN FAMILY
LANCIAN FAMILY
Published: 2016/09/09
Channel: karen tampos
Svizzero no so de lancian
Svizzero no so de lancian
Published: 2017/05/17
Channel: PAULO DYBALA
Warface | Ranked #9
Warface | Ranked #9
Published: 2017/09/07
Channel: Lancian
MILAN LANCIAN
MILAN LANCIAN
Published: 2013/04/21
Channel: Milan Lancian
CELNA LANCIAN FAMILY WE LOVE U ALL UMWAAH
CELNA LANCIAN FAMILY WE LOVE U ALL UMWAAH
Published: 2016/09/13
Channel: karen tampos
Lancian gwapa
Lancian gwapa
Published: 2016/03/17
Channel: christianjames lancian
celna Lancian
celna Lancian
Published: 2017/01/29
Channel: Celna Lancian
Celina Lancian the lonely girl
Celina Lancian the lonely girl
Published: 2015/03/24
Channel: saihane lancian
my little Jeanina Lancian Flores
my little Jeanina Lancian Flores
Published: 2014/03/07
Channel: jean flores
Rallye du Rochois David Lancian Vincent Breynat Lancer évo 6
Rallye du Rochois David Lancian Vincent Breynat Lancer évo 6
Published: 2006/07/13
Channel: vincenzo26320
CSI Equita
CSI Equita'Lyon : Lancian d'Enfer
Published: 2007/11/08
Channel: webstallions
Nellie Lancian Calderon
Nellie Lancian Calderon
Published: 2013/03/16
Channel: nellie calderon
Warface | Ranked #2
Warface | Ranked #2
Published: 2017/06/11
Channel: Lancian
Celna  Lancian.Dela cruz
Celna Lancian.Dela cruz
Published: 2015/03/24
Channel: saihane lancian
12:51 cover by:nellie lancian cadleron
12:51 cover by:nellie lancian cadleron
Published: 2013/03/16
Channel: nellie calderon
Warface | Ranked #5
Warface | Ranked #5
Published: 2017/07/24
Channel: Lancian
M2U01252.MPG lancian grace
M2U01252.MPG lancian grace
Published: 2011/11/29
Channel: lasuertedavao
Sébastien Loeb Rally EVO Lancian Sekoilut
Sébastien Loeb Rally EVO Lancian Sekoilut
Published: 2017/08/12
Channel: Heksu 25
Warface | Ranked #3
Warface | Ranked #3
Published: 2017/06/15
Channel: Lancian
Lancian mix video
Lancian mix video
Published: 2017/04/24
Channel: Celna Lancian
Lanciano-Salernitana:ma che è fatt u lancian?
Lanciano-Salernitana:ma che è fatt u lancian?
Published: 2016/06/05
Channel: Travis1998 Iuzzo
Nellie teach you to do cup song
Nellie teach you to do cup song
Published: 2013/05/11
Channel: Nellie lancian calderon
Fishing using uopilot
Fishing using uopilot
Published: 2017/09/02
Channel: Jan Michael Lancian
Carina ska åka Lancian för första gången.
Carina ska åka Lancian för första gången.
Published: 2017/08/19
Channel: Carina Taxén
Gg stupenda al naviglio!Fra e luca si lancian dal ponte
Gg stupenda al naviglio!Fra e luca si lancian dal ponte
Published: 2008/08/01
Channel: BastardaRossonera
Dirt Rally | "Flugzeugring" Lancia Stratos Top time: 2:47.75
Dirt Rally | "Flugzeugring" Lancia Stratos Top time: 2:47.75
Published: 2017/06/15
Channel: Lancian
Warface | CYBERNETIX vs -SchoolBus- #1
Warface | CYBERNETIX vs -SchoolBus- #1
Published: 2017/06/04
Channel: Lancian
Warface | Ranked #4
Warface | Ranked #4
Published: 2017/07/03
Channel: Lancian
Lancian koneen räjähdys
Lancian koneen räjähdys
Published: 2008/06/23
Channel: tapparinet
Warface | Ranked #7
Warface | Ranked #7
Published: 2017/08/19
Channel: Lancian
Warface | CYBERNETIX vs *H.U.S.A.R.I.A* #2
Warface | CYBERNETIX vs *H.U.S.A.R.I.A* #2
Published: 2016/11/27
Channel: Lancian
cheb aziz aynik mlah lancian de chaoui
cheb aziz aynik mlah lancian de chaoui
Published: 2015/11/02
Channel: Dj Boumediene
Milagre Eucarístico de Lanciano
Milagre Eucarístico de Lanciano
Published: 2010/10/21
Channel: Peregrinações Parceria Obra de Maria e Canção Nova
Warface | CYBERNETIX 4Fun #2
Warface | CYBERNETIX 4Fun #2
Published: 2016/12/11
Channel: Lancian
kamel l
kamel l'ancien
Published: 2008/02/20
Channel: mousstafick enza
Claire-Elie Tenet Meine Lippen Giuditta
Claire-Elie Tenet Meine Lippen Giuditta
Published: 2017/07/15
Channel: Claire TENET
Dirt Rally | "Flugzeugring" Ford Fiesta Top time: 2:38.643
Dirt Rally | "Flugzeugring" Ford Fiesta Top time: 2:38.643
Published: 2017/06/15
Channel: Lancian
Milagre Eucarístico em Kearns-EUA! Hóstia está Sangrando  (#275 - Notícias Assombradas)
Milagre Eucarístico em Kearns-EUA! Hóstia está Sangrando (#275 - Notícias Assombradas)
Published: 2015/12/08
Channel: AssombradO.com.br
17 July 2016
17 July 2016
Published: 2016/07/17
Channel: AnnaRose Lancian
Warface | Trzepiemy Versusy #3
Warface | Trzepiemy Versusy #3
Published: 2017/01/07
Channel: Lancian
Warface | Trzepiemy Versusy #10
Warface | Trzepiemy Versusy #10
Published: 2017/07/26
Channel: Lancian
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WIKIPEDIA ARTICLE

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The Lancian was a North American faunal stage of the Late Cretaceous.

Geology[edit]

Terrestrial sedimentary strata from the Judithian to the Lancian are generally regressive throughout the entire sequence, so the preserved changes in fossil communities represent not only phylogenetic changes but ecological zones from the submontane habitats to near-sea level coastal habitats.[1]

Paleobiogeography[edit]

By the Lancian, the crested hadrosaurs are no longer the dominant inhabitant of any province of western North America; with the exception of Hypacrosaurus.[2] Lehman records three surviving chasmosaurs, Triceratops, Torosaurus and Nedoceratops, with the possibility of the recently discovered Ojoceratops, Regaliceratops and Bravoceratops.[2] It has recently been suggested that Triceratops and Torosaurus may be synonymous, though this is still up for debate. Saurolophine dinosaurs like Edmontosaurus, Kritosaurus, Saurolophus as well as the recently discovered Augustynolophus were some of the known surviving hadrosaurs.[2] Both lack the elaborate ornamentation of their predecessors, the lambeosaurs.[2] The ankylosaurs had been reduced to down to a few species, ankylosaurs like Anodontosaurus and Ankylosaurus, as well as nodosaurs like Denversaurus, Edmontonia and Glyptodontopelta were the only known survivors. In the south the transition to the Lancian is even more dramatic, which Lehman describes as "the abrupt reemergence of a fauna with a superficially "Jurassic" aspect."[2] These faunas are dominated by Alamosaurus and feature abundant Quetzalcoatlus, Bravoceratops and Ojoceratops in Texas.[3]

The extreme changes occurring in the make-up of herbivore communities during the faunal turnover suggests that a change in the ecosystems' flora was "the most immediate cause...though perhaps not the ultimate one."[4] The rapid expansion of land and drying of inland climate accompanying a drop in sea level could explain some of the environmental changes occurring Late Cretaceous western North America.[5] The wetland habitat enjoyed by many dinosaurs would have shrunk and fragmented.[5] Since many species had very limited geographic ranges its plausible that some of the fragments would be smaller than the area needed to support the species.[5] However, there's no direct evidence for the shrinking of wetland environments.[5] Lehman contends that the actual area of coastal lowlands within 150m of the shoreline must have actually increased significantly.[5] Further, dinosaurs that inhabited inland or arid environments were among the most prevalent in the Lancian.[5] The Alamosaurus-Quetzalcoatlus association probably represent semi-arid inland plains.[5] In previous research Jack Horner speculated that a rise in sea level during the Bearpaw Transgression created selective pressure as coastal lowlands were swallowed up the sea, resulting in anagenesis.[6] If the geographic range of some dinosaur species were truly as limited as the fossil record suggests, then a rapid rise and in sea level could cause intense pressure even the event was local.[6] Additionally, a rapid drop in sea level could allow for "rapid colonization by a few dinosaur generalists."[6] The appearance or reappearance of basal neoceratopsians could be explained by immigration from Asia.[6] Dinosaurs like Nodocephalosaurus resembled Asian forms, and some like Saurolophus co-occurred in Asia as well as North America.[6] Potential Asian immigrants were especially common in upland environments.[6] The appearance of Alamosaurus may have represented an immigration event from South America.[6] Some taxa may have co-occurred on both continents, including Kritosaurus and Avisaurus.[6] Alamosaurus appears and achieves dominance in its environment very abruptly.[6] Some scientists speculated that Alamosaurus was an immigrant from Asia.[6] Inhabitants of upland environment are more likely to be endemic than coastal species, and tend to have less of an ability to cross bodies of water.[6] Further, early Cretaceous titanosaurs were already known, so North American potential ancestors for Alamosaurus already existed.[6] Quetzalcoatlus also had precursors in North America and its apparent range expansion may represent the expansion of its preferred habitat rather than an immigration event.[6] Early Cretaceous deposits in North America reveal that basal neoceratopsians were already present on the continent before their apparent reemergence in the Lancian, so an immigration event from Asia is unnecessary to explain their appearance.[6] The major potential immigrants represent archaic forms that probably wouldn't have directly competed with the disappearing forms anyway.[7] Lehman described the evidence for immigration as a driving force in Lancian dinosaur faunal turnovers as "not particularly compelling."[6]

The faunal turnover may be explained by the descent of more primitive forms existing in upland refugia characterized by conifer-dominated flora into areas that were formerly coastal lowlands as the seas retreated and conditions became more arid.[8]

The decline of mammal diversity in Western North America from the Miocene to the present primarily effected large herbivores and occurred over roughly the same length of time as the Late Cretaceous changes, and so may be parallel.[9] They have many commonalities, including the replacement of diverse with single species environments (caribou in the north, bison to the south).[9] The most spectacular and specialized forms became extinct.[9] The mammalian turnover was preceded by an episode of immigration,[9] and was associated with the rapid expansion of terrestrial habitat due to melting glaciers.[9] In the mammalian turnover, the newly emerging dominant fauna were clearly Old World immigrants, the cervids and bovids.[10]

In the southern biome, by Lancian time sauropods had replaced both hadrosaurs and ceratopsians.[11] In the north, both were still present although hadrosaurs were demoted to a "subordinate" role in dinosaur ecosystems.[11] Edmontosaurus was the dominant northern hadrosaurid.[11] At the end of the Cretaceous, most ecosystems were dominated by a single herbivore.[11] The northern biome was dominated by Triceratops and the southern biome by Alamosaurus.[11] This faunal turnover coincides with the Laramide orogeny and the uplift of the central Rockies.[11] Strata exhibit changes in lithology and the direction of paleocurrents, and a severe drop in relative sea level.[11] At the very least, Lehman argues, the altitudinal life zones would shift, and a change in the distribution of vegetation utilized by herbivorous dinosaurs would have probably resulted.[11] By the end of the Judithian, North America had 7.7 million km2 of land area, but by the end of the Lancian it had reached 17.9 million km2, nearly the modern value of 22.5 million km2.[11]

Footnotes[edit]

  1. ^ "Altitudinal and Transcontinental Life Zones," Lehman (2001); pages 312-313.
  2. ^ a b c d e "Lancian Turnover," Lehman (2001); page 317.
  3. ^ "Lancian Turnover," Lehman (2001); pages 317-319.
  4. ^ "What Happened?" Lehman (2001); page 319.
  5. ^ a b c d e f g "Loss of Wetlands Hypothesis," Lehman (2001); page 320.
  6. ^ a b c d e f g h i j k l m n o "Competition from Invaders Hypothesis," Lehman (2001); page 321.
  7. ^ "Competition from Invaders Hypothesis," Lehman (2001); pages 321-322.
  8. ^ "Descent from the Highlands Hypothesis," Lehman (2001); page 322.
  9. ^ a b c d e "A Recent Analog?" Lehman (2001); page 323.
  10. ^ "A Recent Analog?" Lehman (2001); pages 323-324.
  11. ^ a b c d e f g h i "Conclusions," Lehman (2001); page 324.

References[edit]

  • Cifelli, Richard L.; Eberle, Jaelyn J.; Lofgren, Donald L.; Lillegraven, Jason A.; Clemens, William A. (2012). "Mammalian Biochronology of the Latest Cretaceous". In Woodburne, Michael O. Late Cretaceous and Cenozoic mammals of North America biostratigraphy and geochronology. New York: Columbia University Press. ISBN 9780231503785. 
  • Kielan-Jaworowska, Zofia; Cifelli,, Richard L.; Luo, Zhe-Xi (2004). Mammals from the age of dinosaurs origins, evolution, and structure. New York: Columbia University. pp. 19–108. ISBN 9780231509275. 
  • Lehman, T. M., 2001, Late Cretaceous dinosaur provinciality: In: Mesozoic Vertebrate Life, edited by Tanke, D. H., and Carpenter, K., Indiana University Press, pp. 310–328.
  • Lucas, Spencer G, ed. (1991). "Dinosaurs and Mesozoic biochronology". Modern Geology. 16: 127–138.  Reprinted in Halstead, L.B. (1991). Dinosaur studies : commemorating the 150th anniversary of Richard Owen's Dinosauria. Philadelphia: Gordon and Breach. ISBN 9782881248337. 

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