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TANYCOLAGREUS  MAX LEVEL 40 - NEW VIP DINO Jurassic World The Game
TANYCOLAGREUS MAX LEVEL 40 - NEW VIP DINO Jurassic World The Game
Published: 2016/07/19
Channel: Gam1ngNinja
TANYCOLAGREUS MAX LVL 40! -Jurassic World The Game - *NEW DINO* HD
TANYCOLAGREUS MAX LVL 40! -Jurassic World The Game - *NEW DINO* HD
Published: 2016/07/01
Channel: LPGozzzi
TANYCOLAGREUS VS TANYCOLAGREUS ?! VIP DINOSAUR ?! |Jurassic World The Game|Ep 130
TANYCOLAGREUS VS TANYCOLAGREUS ?! VIP DINOSAUR ?! |Jurassic World The Game|Ep 130
Published: 2016/10/03
Channel: Vertico
Tanycolagreus and Mastodonsaurus Max Level - Jurassic World The Game
Tanycolagreus and Mastodonsaurus Max Level - Jurassic World The Game
Published: 2017/02/27
Channel: Alex Greenland
TANYCOLAGREUS DINOSSAURO MAIS FORTE! - Jurassic World - O Jogo - Ep 42
TANYCOLAGREUS DINOSSAURO MAIS FORTE! - Jurassic World - O Jogo - Ep 42
Published: 2016/07/20
Channel: Piu
Tanycolagreus - Video Learning - WizScience.com
Tanycolagreus - Video Learning - WizScience.com
Published: 2015/08/05
Channel: Wiz Science™
GANHEI UM TANYCOLAGREUS PACOTE VIP! - Jurassic World - O Jogo - Ep 139
GANHEI UM TANYCOLAGREUS PACOTE VIP! - Jurassic World - O Jogo - Ep 139
Published: 2016/12/13
Channel: Piu
TANYCOLAGREUS Maxed - Jurassic World The Game - VIP Dinosaur
TANYCOLAGREUS Maxed - Jurassic World The Game - VIP Dinosaur
Published: 2016/07/18
Channel: BorkTheBold
New Tanycolagreus STK
New Tanycolagreus STK
Published: 2016/12/28
Channel: Neovenator Rex
TANYCOLAGREUS in Jurassic World: The Game - Part 4
TANYCOLAGREUS in Jurassic World: The Game - Part 4
Published: 2016/10/14
Channel: Game24h.com
VIP TANYCOLAGREUS Vs SARCOSUCHUS!! || Jurassic World - The Game Gameplay #13
VIP TANYCOLAGREUS Vs SARCOSUCHUS!! || Jurassic World - The Game Gameplay #13
Published: 2016/08/22
Channel: iGameplay1224
TANYCOLAGREUS NO LEVEL 30! - Jurassic World - O Jogo - Ep 177
TANYCOLAGREUS NO LEVEL 30! - Jurassic World - O Jogo - Ep 177
Published: 2017/02/07
Channel: Piu
TANYCOLAGREUS PACOTE VIP 30000 PONTOS! - Jurassic World - O Jogo - Ep 40
TANYCOLAGREUS PACOTE VIP 30000 PONTOS! - Jurassic World - O Jogo - Ep 40
Published: 2016/07/19
Channel: Piu
Evoluindo Tanycolagreus Vip + Eventos &  Melhor Time do YouTube ptBR JURASSIC WORLD O JOGO THE GAME
Evoluindo Tanycolagreus Vip + Eventos & Melhor Time do YouTube ptBR JURASSIC WORLD O JOGO THE GAME
Published: 2017/06/15
Channel: Elite-AlexTracer
VIP TANYCOLAGREUS MAX LEVEL FEEDING | WORLD BOSS T.Rex Omega 09 - Jurassic World The Game
VIP TANYCOLAGREUS MAX LEVEL FEEDING | WORLD BOSS T.Rex Omega 09 - Jurassic World The Game
Published: 2016/07/01
Channel: PlayCow.com
Jurassic World: The Game - Player Duel Battle 639 - TANYCOLAGREUS!
Jurassic World: The Game - Player Duel Battle 639 - TANYCOLAGREUS!
Published: 2016/09/23
Channel: EmiPivarchuk10
TANYCOLAGREUS nivel 40
TANYCOLAGREUS nivel 40
Published: 2016/07/01
Channel: DINO GAMES
Tanycolagreus sounds
Tanycolagreus sounds
Published: 2016/07/05
Channel: Cj Magruder
Jurassic World: Das Spiel #102 Tanycolagreus Lvl. 40 & Protostega!!  [60FPS/HD] | Marcel
Jurassic World: Das Spiel #102 Tanycolagreus Lvl. 40 & Protostega!! [60FPS/HD] | Marcel
Published: 2016/11/25
Channel: TheBrawlyBeaters
TANYCOLAGREUS & PRESTOSSUCO +Novidades Vip + Batalhas - Jurassic World: O Jogo
TANYCOLAGREUS & PRESTOSSUCO +Novidades Vip + Batalhas - Jurassic World: O Jogo
Published: 2016/09/17
Channel: Elite-AlexTracer
Jurassic World - The Game || VIP Max Level TANYCOLAGREUS Vs TYRANNOSAURUS REX Gameplay HD #8
Jurassic World - The Game || VIP Max Level TANYCOLAGREUS Vs TYRANNOSAURUS REX Gameplay HD #8
Published: 2016/08/10
Channel: iGameplay1224
New  Jurassic World The Game  VIP Battles Episode 114 Tanycolagreus Vs Indominus Dinosaurs WD Toys
New Jurassic World The Game VIP Battles Episode 114 Tanycolagreus Vs Indominus Dinosaurs WD Toys
Published: 2016/10/05
Channel: WDTechToys
Jurassic World: The Game - SOLID GOLD PACK - TANYCOLAGREUS - Level 20 Evolution + 1st Battle
Jurassic World: The Game - SOLID GOLD PACK - TANYCOLAGREUS - Level 20 Evolution + 1st Battle
Published: 2016/10/08
Channel: EmiPivarchuk10
Jurassic World - The Game || TANYCOLAGREUS Vs INDOMINUS REX - Gameplay Full HD #11
Jurassic World - The Game || TANYCOLAGREUS Vs INDOMINUS REX - Gameplay Full HD #11
Published: 2016/08/17
Channel: iGameplay1224
TANYCOLAGREUS Max LV 40 V.I.P Pack - Jurassic World The Game
TANYCOLAGREUS Max LV 40 V.I.P Pack - Jurassic World The Game
Published: 2016/10/06
Channel: The GAMES
New Jurassic World The Game VIP Battles Episode 116 Tanycolagreus Vs Indominus Dinosaurs WD Toys
New Jurassic World The Game VIP Battles Episode 116 Tanycolagreus Vs Indominus Dinosaurs WD Toys
Published: 2016/10/07
Channel: WDTechToys
Tanycolagreus Pack & The Colossus of Canada!!!-Jurassic World:The Game Ep. #17
Tanycolagreus Pack & The Colossus of Canada!!!-Jurassic World:The Game Ep. #17
Published: 2016/12/31
Channel: Terex's Planet
Tanycolagreus max level 40+card pack||Jurassic World the Game
Tanycolagreus max level 40+card pack||Jurassic World the Game
Published: 2017/05/19
Channel: Alex Gamer
Tanycolagreus Level 20!!!-Jurassic World:The Game Ep. #19
Tanycolagreus Level 20!!!-Jurassic World:The Game Ep. #19
Published: 2017/01/05
Channel: Terex's Planet
Jurassic World: The Game - Battle Stage 63: Scaphognathus - TANYCOLAGREUS Level 30 Debut!
Jurassic World: The Game - Battle Stage 63: Scaphognathus - TANYCOLAGREUS Level 30 Debut!
Published: 2016/11/13
Channel: EmiPivarchuk10
Jurassic World - The Game ► Tanycolagreus | EP 97
Jurassic World - The Game ► Tanycolagreus | EP 97
Published: 2017/02/08
Channel: Wero
TANYCOLAGREUS NEW VIP LEGENDARY DINOSAUR Max Level 40 - Jurassic World the Game - Live Arena 2016
TANYCOLAGREUS NEW VIP LEGENDARY DINOSAUR Max Level 40 - Jurassic World the Game - Live Arena 2016
Published: 2016/07/19
Channel: VBGAMING11
Jurassic World: The Game - Player Duel Battle 660 - TANYCOLAGREUS!
Jurassic World: The Game - Player Duel Battle 660 - TANYCOLAGREUS!
Published: 2016/09/26
Channel: EmiPivarchuk10
Getting Tanycolagreus to LEVEL 40!| Jurassic World The Game
Getting Tanycolagreus to LEVEL 40!| Jurassic World The Game
Published: 2016/09/17
Channel: #Absolutely No Theme#
Jurassic World - Max Level 40 VIP Legendary Tanycolagreus
Jurassic World - Max Level 40 VIP Legendary Tanycolagreus
Published: 2016/09/17
Channel: BagoyeeGaming
Jurassic World: The Game - Player Duel Battle 746 - TANYCOLAGREUS
Jurassic World: The Game - Player Duel Battle 746 - TANYCOLAGREUS
Published: 2016/10/13
Channel: EmiPivarchuk10
Jurassic World: The Game - "VIP Only Event" 4 - Full Battle - TANYCOLAGREUS And Indominus Rex
Jurassic World: The Game - "VIP Only Event" 4 - Full Battle - TANYCOLAGREUS And Indominus Rex
Published: 2016/07/28
Channel: EmiPivarchuk10
Jurassic World: The Game - Player Duel Battle 770 - TANYCOLAGREUS And PTERODACTYLUS VS PRESTOSUCHUS!
Jurassic World: The Game - Player Duel Battle 770 - TANYCOLAGREUS And PTERODACTYLUS VS PRESTOSUCHUS!
Published: 2016/10/22
Channel: EmiPivarchuk10
Jurassic World: The Game - SPECIAL - My 2nd VIP Dinosaur: TANYCOLAGREUS!
Jurassic World: The Game - SPECIAL - My 2nd VIP Dinosaur: TANYCOLAGREUS!
Published: 2016/09/19
Channel: EmiPivarchuk10
Ep 44 || battles + tanycolagreus level 20 - Jurassic world the game!
Ep 44 || battles + tanycolagreus level 20 - Jurassic world the game!
Published: 2016/12/10
Channel: The KillaRex
Jurassic World: The Game - Player Duel Battle 573 - TANYCOLAGREUS!
Jurassic World: The Game - Player Duel Battle 573 - TANYCOLAGREUS!
Published: 2016/08/18
Channel: EmiPivarchuk10
Jurassic World: The game - Tanycolagreus MAX LVL
Jurassic World: The game - Tanycolagreus MAX LVL
Published: 2016/07/15
Channel: OhItsAndrew
TANYCOLAGREUS NEW LEGENDARY DINOSAUR VIP UPDATE Max Level 40 - Jurassic World - Epic Gameplay 2016
TANYCOLAGREUS NEW LEGENDARY DINOSAUR VIP UPDATE Max Level 40 - Jurassic World - Epic Gameplay 2016
Published: 2016/06/30
Channel: VBGAMING11
Evoluindo VIP Tanycolagreus do nível 1 ao 30 JURASSIC WORLD O JOGO
Evoluindo VIP Tanycolagreus do nível 1 ao 30 JURASSIC WORLD O JOGO
Published: 2016/12/04
Channel: Elite-AlexTracer
Jurassic World: The Game - Player Duel Battle 708 - TANYCOLAGREUS VS TANYCOLAGREUS!
Jurassic World: The Game - Player Duel Battle 708 - TANYCOLAGREUS VS TANYCOLAGREUS!
Published: 2016/10/05
Channel: EmiPivarchuk10
Jurassic World - The Game || TANYCOLAGREUS Vs SPINOSAURUS Gameplay Full HD #11
Jurassic World - The Game || TANYCOLAGREUS Vs SPINOSAURUS Gameplay Full HD #11
Published: 2016/08/16
Channel: iGameplay1224
TANYCOLAGREUS MAX LEVEL 40 | JURASSIC WORLD
TANYCOLAGREUS MAX LEVEL 40 | JURASSIC WORLD
Published: 2016/09/20
Channel: Nguyên Vương
TANYCOLAGREUS | Jurassic World - El Juego | #37 | Gameplay Español
TANYCOLAGREUS | Jurassic World - El Juego | #37 | Gameplay Español
Published: 2017/03/12
Channel: BellsproutArmy
TANYCOLAGREUS NEW VIP LEGENDARY DINOSAUR UPDATE Max Level 40 - Jurassic World - Omega 09 Battle 2016
TANYCOLAGREUS NEW VIP LEGENDARY DINOSAUR UPDATE Max Level 40 - Jurassic World - Omega 09 Battle 2016
Published: 2016/07/01
Channel: VBGAMING11
WORLD EVENT BOSS - OMEGA 09 LEVEL 04 Stop TYRANNOSAURUS - Tanycolagreus,Apatosaurus New VIP UPDATE
WORLD EVENT BOSS - OMEGA 09 LEVEL 04 Stop TYRANNOSAURUS - Tanycolagreus,Apatosaurus New VIP UPDATE
Published: 2016/06/30
Channel: VBGAMING11
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WIKIPEDIA ARTICLE

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Tanycolagreus
Temporal range: Late Jurassic, 153–150 Ma
Tanycolagreus.jpg
Tanycolagreus holotype, North American Museum of Ancient Life.
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Dinosauria
Order: Saurischia
Suborder: Theropoda
Genus: Tanycolagreus
Carpenter et al., 2005
Species: T. topwilsoni
Binomial name
Tanycolagreus topwilsoni
Carpenter et al., 2005

Tanycolagreus is a genus of coelurosaurian theropod from the Late Jurassic of North America.

Description[edit]

Restoration of Tanycolagreus topwilsoni

Carpenter et al. (2005, pp. 43–44) determined that the holotype of Tanycolagreus represents a subadult individual which measured approximately 3.3 meters (11 ft) long in life. However, one of the referred fossils, the premaxilla from the Cleveland-Lloyd Quarry, would have belonged to a larger individual, measuring 4 meters (13 ft) long. In 2010 Gregory S. Paul estimated the weight of a four metres long animal at hundred twenty kilogrammes.[1] It cannot be determined whether or not the Cleveland-Lloyd specimen represents a fully mature adult, so the upper size limit for the taxon remains unknown.

The head of Tanycolagreus is large, elongated and rectangular in profile due to a blunt snout. The leg is rather long and lightly built.[1]

Restored skeleton

Carpenter et al. (2005; pp. 27 & 29) diagnosed Tanycolagreus topwilsoni as follows: "Medium-sized tetanuran having short, deep-bodied premaxilla pierced by narial foramen at base of nasal process, orbital process on postorbital, T-shaped quadratojugal, centrodiapophyseal lamina on dorsals. Differs from Coelurus in the absence of pleurocoel on anterior dorsals; posterior caudal prezygapophyses elongated to one-third centrum length, rather than short; straight, rather than sigmoidal, humeral shaft; bowed, rather than straight, radius; flat-bottomed rather than arced pubic foot; straight rather than sigmoidal femoral shaft; metatarsal length subequal to humeral length, rather than 1.75 times humeral length. Differs from Ornitholestes in straight anterior margin of premaxilla, rather than rounded; T-shaped rather than L-shaped quadratojugal; elongate neural spine; posterior caudal prezygapophyses only one-third centrum length, rather than one-half centrum length; bowed, slender radius, rather than straight, robust radius".[2]

The single premaxillary tooth preserved with the holotype is badly damaged, but does exhibit the asymmetrical cross-section typical in theropod teeth; the cheek teeth are too poorly preserved to show any detail. In the foot the second toe is slightly hyperextendable but does not carry an enlarged claw.[2]

Discovery and naming[edit]

Restored skull cast

In 1995 Western Paleontological Laboratories, Inc. uncovered the partial skeleton of a small theropod at the Bone Cabin Quarry West locality, Albany County, Wyoming, from the Salt Wash Member of the Morrison Formation, dating to the Oxfordian-Tithonian. At first the find was considered to be a specimen of Coelurus[3] but subsequent study indicated it represented a species new to science, that in 2001 was announced to be named Tanycolagreus topwilsoni. It was actually named and described by Kenneth Carpenter, Clifford Miles and Karen Cloward in 2005. The etymology of the generic name Tanycolagreus, suggested by Ben Creisler, is based upon the greater length of its forelimbs and hindlimbs compared to Coelurus. It is derived from the Greek prefix τανυ~, tany~: 'long, stretched out', κῶλον, kolon: 'limb' and ἀγρεύς, agreus: 'hunter'. The specific name honours George Eugene "Top" Wilson, the father of a benefactor financially supporting the scientific research.[2]

The fossil, holotype TPII 2000-09-29, was donated to science by an anonymous benefactor. It is part of the collection of Thanksgiving Point Institute, Inc. and displayed in the North American Museum of Ancient Life at Lehi, Utah. It includes an incomplete skull and mandible (lower jaws) and much of the postcranial skeleton, i.e. the parts behind the head. The skull of Tanycolagreus is less well known than its postcranial anatomy, and only the following elements have been found: left nasal, left lacrimal, left premaxilla and one premaxillary tooth, left postorbital, left quadratojugal, incomplete left squamosal, right quadrate, right splenial, left articular, and two cheek teeth. A paratype has been assigned to the species: specimen AMNH 587 consisting of an incomplete hand also collected from Bone Cabin Quarry and originally in 1903 by Henry Fairfield Osborn referred to Ornitholestes hermanni.[4] Two other fossils have been referred to Tanycolagreus: UUVP 2999, a premaxilla, originally in 1974 referred to Stokesosaurus clevelandi,[5] from the Cleveland-Lloyd Quarry of Utah; and USNM 5737, a pair of distal pubes from Colorado earlier in 1920 by Charles Whitney Gilmore referred to Coelurus.[6] These specimens are from the later Brushy Basin Member.[2]

Tanycolagreus is present in stratigraphic zone 2 of the Morrison. Remains possibly referrable to Stokesosaurus have been recovered from stratigraphic zone 5 of the Morrison Formation.[7] A life restoration of Tanycolagreus is also on display at the North American Museum of Ancient Life, where it is portrayed as preying upon a small ornithischian dinosaur, Othnielia rex.

Classification[edit]

Carpenter e.a. originally assigned Tanycolagreus to the Coeluridae.[2] Carpenter et alii (2005, p. 44) state that, of the other known Morrison theropods, this genus most closely resembles Coelurus, though it retains more "primitive" features. A detailed phylogenetic analysis in 2007 by Philip Senter including Tanycolagreus showed it had a basal position in the Tyrannosauroidea.[8] Later analyses indicated a basal position in the Coelurosauria.[9]

References[edit]

  1. ^ a b Paul, G.S., 2010, The Princeton Field Guide to Dinosaurs, Princeton University Press p. 124
  2. ^ a b c d e Carpenter, K., Miles, C., and Cloward, K. (2005). "New small theropod from the Upper Jurassic Morrison Formation of Wyoming." in Carpenter, K. 2005. The Carnivorous Dinosaurs, Indiana University Press: 23-48
  3. ^ Miles, C.A., Carpenter K. and Cloward, K.C., 1998, "A new skeleton of Coelurus fragilis from the Morrison Formation of Wyoming", Journal of Vertebrate Paleontology 18(3): 64A
  4. ^ Osborn, Henry Fairfield, 1903, "Ornitholestes hermanni, a new compsognathoid dinosaur from the Upper Jurassic", Bulletin of the American Museum of Natural History 19(12): 459–464
  5. ^ Madsen, J., 1974, "A new theropod dinosaur from the Upper Jurassic of Utah", Journal of Paleontology, 48: 27-31
  6. ^ Gilmore, C.W., 1920, "Osteology of the carnivorous Dinosauria in the United States National Museum, with special reference to the genera Antrodemus (Allosaurus) and Ceratosaurus", Bulletin of the United States National Museum, 110: 1-154
  7. ^ Foster, J. (2007). "Appendix." Jurassic West: The Dinosaurs of the Morrison Formation and Their World. Indiana University Press. pp. 327-329
  8. ^ Senter, P. (2007). "A new look at the phylogeny of Coelurosauria (Dinosauria: Theropoda)", Journal of Systematic Palaeontology, 5(4): 429-463
  9. ^ O.W.M. Rauhut, A.C. Milner, and S. Moore-Fay, 2010, "Cranial osteology and phylogenetic position of the theropod dinosaur Proceratosaurus bradleyi (Woodward, 1910) from the Middle Jurassic of England", Zoological Journal of the Linnean Society, 158(1): 155-195

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