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Wild Monitor Lizard - Varanidae - Lagarto gigante - Singapore
Wild Monitor Lizard - Varanidae - Lagarto gigante - Singapore
Published: 2015/03/02
Channel: Wild Animals
Varanidae - Video Learning - WizScience.com
Varanidae - Video Learning - WizScience.com
Published: 2015/09/17
Channel: Wiz Science™
Varanidae - American Boy
Varanidae - American Boy
Published: 2015/08/21
Channel: Attitude Music
Monitor Lizards Do Battle! (Lumphini Park - Bangkok, Thailand)
Monitor Lizards Do Battle! (Lumphini Park - Bangkok, Thailand)
Published: 2009/02/01
Channel: Nicholas Snow
Dragão de komodo comendo um veado vivo  impressionante animal com veneno mortal
Dragão de komodo comendo um veado vivo impressionante animal com veneno mortal
Published: 2017/07/30
Channel: CANAL ANIMAL NEWSPAPER EM ALTA
Варан играется со шлангом
Варан играется со шлангом
Published: 2015/12/30
Channel: Частный экзотариум
Monitor lizard in the wild
Monitor lizard in the wild
Published: 2010/09/01
Channel: WildFilmsIndia
IndianMonitorLizard गोह( ઘો )_घोरपड
IndianMonitorLizard गोह( ઘો )_घोरपड
Published: 2017/05/25
Channel: DarSnake - The Reptile Rescuer
Капский варан
Капский варан
Published: 2016/04/16
Channel: Частный экзотариум
Komodo!The Deadly Encounter With A Monster Dragon.
Komodo!The Deadly Encounter With A Monster Dragon.
Published: 2017/08/07
Channel: jit mukherjee
Bắt và nướng kỳ đà khủng | Catch and Cook Varanidae in the Jungle
Bắt và nướng kỳ đà khủng | Catch and Cook Varanidae in the Jungle
Published: 2016/06/05
Channel: Jet paso
Biawak - Bayawak - Menyawak - Nyambik - Berekai - Monitor lizard - Goanna - Varanidae
Biawak - Bayawak - Menyawak - Nyambik - Berekai - Monitor lizard - Goanna - Varanidae
Published: 2014/10/15
Channel: Ispinardi Harjanto
Varaan op vakantie in Maleisië (Varanidae)
Varaan op vakantie in Maleisië (Varanidae)
Published: 2011/08/10
Channel: tvfreak7777
Papua lizard أقصى चरम ekstrim
Papua lizard أقصى चरम ekstrim
Published: 2012/09/19
Channel: Mieke Peters
Fishing Adventure Thailand by Nopbkk - Lucky Rat vs Varanidae
Fishing Adventure Thailand by Nopbkk - Lucky Rat vs Varanidae
Published: 2014/02/20
Channel: Nop Karnjanabhan
Komodo dragon (Varanus Komodoensis)
Komodo dragon (Varanus Komodoensis)
Published: 2010/05/01
Channel: Animal education
Spiderman: Ataxia | Episode 7 "Varanidae"
Spiderman: Ataxia | Episode 7 "Varanidae"
Published: 2016/01/10
Channel: EV3Gurl
Varanidae - Remy [MEOW!]
Varanidae - Remy [MEOW!]
Published: 2015/08/23
Channel: Foxty Music
Vast desert monitor in Reptilium Landau Familie Varanidae
Vast desert monitor in Reptilium Landau Familie Varanidae
Published: 2013/09/23
Channel: Prof_LOL
CIUM - film grozy. Waran Stepowy
CIUM - film grozy. Waran Stepowy
Published: 2015/09/16
Channel: DzonyDwururka
Varanidae - Rainbows | ER Free Release
Varanidae - Rainbows | ER Free Release
Published: 2015/08/23
Channel: ELECTRONIC RAVERS
Varudhini Parinayam - Watch Full Episode 1 of 5th August 2013
Varudhini Parinayam - Watch Full Episode 1 of 5th August 2013
Published: 2013/08/23
Channel: zeetvtelugu
Pertarungan Biawak Vs Cicak - Varanidae VS Gekkonidae
Pertarungan Biawak Vs Cicak - Varanidae VS Gekkonidae
Published: 2016/12/18
Channel: REYDI OKTAVIAN
TIJC2014 Varanidae, Dan Phillips : Silpakon University Jazz Faculty
TIJC2014 Varanidae, Dan Phillips : Silpakon University Jazz Faculty
Published: 2014/09/25
Channel: Thailand International Jazz Conference TIJC
Dragão de Komodo, O Lagarto Gigante da Indonésia
Dragão de Komodo, O Lagarto Gigante da Indonésia
Published: 2009/08/15
Channel: Coongo
How to Pronounce Varanidae
How to Pronounce Varanidae
Published: 2014/09/23
Channel: Emma Saying
Monitor lizard विषखोपड़ा Wildlife & Reptile Rescue Release Video.
Monitor lizard विषखोपड़ा Wildlife & Reptile Rescue Release Video.
Published: 2017/07/07
Channel: DarSnake - The Reptile Rescuer
Lace monitor (Varanus varius)
Lace monitor (Varanus varius)
Published: 2011/02/21
Channel: Animal education
Dragão de komodo vs cobra       ( Dragão de komodo e conhecido como o terror das cobras )
Dragão de komodo vs cobra ( Dragão de komodo e conhecido como o terror das cobras )
Published: 2017/07/27
Channel: CANAL ANIMAL NEWSPAPER EM ALTA
How to Pronounce Varanidae - Correct Pronunciation
How to Pronounce Varanidae - Correct Pronunciation
Published: 2017/02/11
Channel: Correct Pronunciation
How to pronounce Varanidae
How to pronounce Varanidae
Published: 2017/01/31
Channel: How to pronounce
IndianMonitorLizard गोह ( ઘો )
IndianMonitorLizard गोह ( ઘો )
Published: 2017/04/21
Channel: DarSnake - The Reptile Rescuer
Szavanna varánusz etetése (Varanus exanthematicus)
Szavanna varánusz etetése (Varanus exanthematicus)
Published: 2009/10/24
Channel: TheVaranidae
Komodo Dragon
Komodo Dragon
Published: 2010/01/13
Channel: Crinale819
Komodo Dragon - description
Komodo Dragon - description
Published: 2017/07/04
Channel: Multicultural Videos
Komodo Dragon - Largest LIZARD on Planet
Komodo Dragon - Largest LIZARD on Planet
Published: 2017/09/06
Channel: JAKHD
What Komoodo Varaan brings to the team
What Komoodo Varaan brings to the team
Published: 2016/11/20
Channel: jouki jou
Dani Varan -Sincera iubire
Dani Varan -Sincera iubire
Published: 2015/09/20
Channel: Mesagerul Cinzecimii
Komodo Islands - Nature Conservancy of Komodo
Komodo Islands - Nature Conservancy of Komodo
Published: 2012/11/07
Channel: Alvin Alexander
Komodo Dragon La Zoo
Komodo Dragon La Zoo
Published: 2010/06/09
Channel: Ezra Kamer
Черный древесный варан пытается выхватить кусочек мяса из кулачка))
Черный древесный варан пытается выхватить кусочек мяса из кулачка))
Published: 2016/01/01
Channel: Частный экзотариум
Đại Chiến Săn Mồi Đẫm Máu của Rồng KOMODO - Komodo dragon war hunting prey
Đại Chiến Săn Mồi Đẫm Máu của Rồng KOMODO - Komodo dragon war hunting prey
Published: 2017/05/31
Channel: Fishing Thích Câu Cá
Banciu: Varanul ii da in judecata pe cei care l-au condamnat
Banciu: Varanul ii da in judecata pe cei care l-au condamnat
Published: 2014/09/30
Channel: B1TVChannel
Dancing of Komodo Dragon on manas  whistle  9213534506 poisonous lizard
Dancing of Komodo Dragon on manas whistle 9213534506 poisonous lizard
Published: 2016/09/11
Channel: Manas Ranjan Mahapatra
Monitor lizard in hole
Monitor lizard in hole
Published: 2013/04/27
Channel: WildFilmsIndia
Varano del Nilo Comiendo (Varanus niloticus)
Varano del Nilo Comiendo (Varanus niloticus)
Published: 2011/07/18
Channel: AntonioRockSnake
Кормление варана
Кормление варана
Published: 2015/09/10
Channel: Частный экзотариум
Komodo Dragon Fighting in Bali Safari Marine Park
Komodo Dragon Fighting in Bali Safari Marine Park
Published: 2017/04/04
Channel: Saurabh Haldiya
Komodo Dragon Mating- Animal Breeding
Komodo Dragon Mating- Animal Breeding
Published: 2015/04/21
Channel: Reptiles Mating
Waran nilowy (Varanus niloticus) i jajko
Waran nilowy (Varanus niloticus) i jajko
Published: 2009/10/03
Channel: epiphone911007
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WIKIPEDIA ARTICLE

From Wikipedia, the free encyclopedia
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Varanids
Temporal range:
Late Cretaceous - Holocene, 80–0 Ma
KomodoDragonRinca1.jpg
Komodo dragon (Varanus komodoensis)
Varanus exanthematicus in the wild.jpg
Savannah monitor (Varanus exanthematicus)
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Squamata
Superfamily: Varanoidea
Family: Varanidae
Merrem, 1820
Genera

See text

The Varanidae (varans) are a family of lizards of the superfamily Varanoidea. The family, a group of carnivorous and frugivorous lizards,[1] includes the extinct Megalania (the largest known lizard), the Komodo dragon (the largest living lizard), and the crocodile monitor. The Varanidae contain the living genus Varanus and a number of extinct taxa. Their closest living relatives are the anguid and helodermatid lizards.[2]

Taxonomy[edit]

Varanidae was defined by Estes, de Queiroz and Gauthier (1988) as the clade containing the most recent common ancestor of Lanthanotus and Varanus and all of its descendants.[3] A similar definition was formulated by Conrad et al. (2008), who defined Varanidae as the clade containing Varanus varius, Lanthanotus borneensis, and all descendants of their last common ancestor.[4] Using one of these definitions leads to the inclusion of the earless monitor lizard (Lanthanotus borneensis) in the family Varanidae.

Lee (1997) created a different definition of Varanidae, defining it as the clade containing Varanus and all taxa more closely related to Varanus than to Lanthanotus;[5][6] this definition explicitly excludes the earless monitor lizard from Varanidae. Whether Lanthanotus borneensis is included in or excluded from Varanidae depends on the author: for example, Vidal et al. (2012) classify the earless monitor lizard as a member of a separate family Lanthanotidae,[7] while Gauthier et al. (2012) classify it as a member of Varanidae.[8]

Genera[edit]

The extinct Saniwa ensidens
Genera marked with are extinct

Genera usually included in Varanidae (under subfamily Varaninae according to Conrad et al., 2008)):

Genera sometimes included in Varanidae (under subfamily Lanthanotinae according to Conrad et al., 2008). Treated as under the separate family Lanthanotidae by other authors:

Basal varanoids:

Formerly included in Varanidae:

Phylogeny[edit]

Below is a cladogram from Conrad et al. (2008) that shows relationships within Varanoidea:[12]

Varanoidea 

Mosasaurs




"Saniwa" feisti




Necrosaurus





Saniwides



Telmasaurus



 Varanidae 
 Lanthanotinae 

Aiolosaurus




Lanthanotus



Cherminotus




 Varaninae 

Ovoo




Saniwa



Varanus









Biology[edit]

Gray's monitor (Varanus olivaceus) is a tree-dwelling varanid from the Philippines that primarily feeds on fruit

Monitor lizards are reputed to be among the most intelligent lizards. Most species forage widely and have large home ranges,[13] and many have high stamina.[14] Although most species are carnivorous, three arboreal species in the Philippines (Varanus olivaceus, Varanus mabitang, and Varanus bitatawa) are primarily frugivores.[1][15] Among species of living varanids, the limbs show positive allometry, being larger in larger-bodied species, although the feet become smaller as compared with the lengths of the other limb segments.[16]

Varanids possess unidirectional pulmonary airflow, including airsacs akin to those of birds.[17]

See also[edit]

References[edit]

  1. ^ a b Welton, L. J.; Siler, C. D.; Bennett, D.; Diesmos, A.; Duya, M. R.; Dugay, R.; Rico, E. L. B.; Van Weerd, M.; Brown, R. M. (2010). "A spectacular new Philippine monitor lizard reveals a hidden biogeographic boundary and a novel flagship species for conservation". Biology Letters. 6 (5): 654–658. ISSN 1744-9561. PMC 2936141Freely accessible. PMID 20375042. doi:10.1098/rsbl.2010.0119. 
  2. ^ Fry, B.G.; Vidal, N; Norman J.A.; Vonk F.J.; Scheib, H.; Ramjan S.F.R; Kuruppu S.; Fung, K.; Hedges, B.; Richardson M.K.; Hodgson, W.C.; Ignjatovic, V.; Summerhays, R.; Kochva, E. (February 2006). "Early evolution of the venom system in lizards and snakes" (PDF). Nature. 439 (7076): 584–588. PMID 16292255. doi:10.1038/nature04328. 
  3. ^ Estes, Richard; Kevin de Queiroz; Jacques Gauthier (1988). "Phylogenetic Relationships within Squamata". In Richard J. Estes. Phylogenetic Relationships of the Lizard Families: Essays Commemorating Charles L. Camp. Stanford University Press. p. 166. ISBN 9780804714358. 
  4. ^ a b Conrad J. (2008). "Phylogeny and systematics of Squamata (Reptilia) based on morphology". Bulletin of the American Museum of Natural History. 310: 1–182. doi:10.1206/310.1. 
  5. ^ Lee MSY (1997-01-29). "The phylogeny of varanoid lizards and the affinities of snakes". Philos Trans R Soc Lond B Biol Sci. 352 (1349): 53–91. PMC 1691912Freely accessible. doi:10.1098/rstb.1997.0005. 
  6. ^ Michael S.Y Lee (2005). "Molecular evidence and marine snake origins". Biology Letters. 1 (2): 227–230. PMC 1626205Freely accessible. PMID 17148173. doi:10.1098/rsbl.2004.0282. 
  7. ^ Nicolas Vidal, Julie Marin, Julia Sassi, Fabia U. Battistuzzi, Steve Donnellan, Alison J. Fitch, Bryan G. Fry, Freek J. Vonk, Ricardo C. Rodriguez de la Vega, Arnaud Couloux and S. Blair Hedges (2012). "Molecular evidence for an Asian origin of monitor lizards followed by Tertiary dispersals to Africa and Australasia". Biology Letters. 8 (5): 853–855. PMC 3441001Freely accessible. PMID 22809723. doi:10.1098/rsbl.2012.0460. 
  8. ^ Jacques A. Gauthier, Maureen Kearney, Jessica Anderson Maisano, Olivier Rieppel, Adam D.B. Behlke; Kearney; Maisano; Rieppel; Behlke (2012). "Assembling the Squamate Tree of Life: Perspectives from the Phenotype and the Fossil Record". Bulletin of the Peabody Museum of Natural History. 53 (1): 3–308. doi:10.3374/014.053.0101. 
  9. ^ Massimo Delfino, Jean-Claude Rage, Arnau Bolet and David M. Alba (2013). "Synonymization of the Miocene varanid lizard Iberovaranus Hoffstetter, 1969 with Varanus Merrem, 1820". Acta Palaeontologica Polonica. in press. doi:10.4202/app.2012.0025. 
  10. ^ a b c Conrad, JL; Balcarcel, AM; Mehling, CM (2012). "Earliest Example of a Giant Monitor Lizard (Varanus, Varanidae, Squamata)". PLoS ONE. 7 (8): e41767. PMC 3416840Freely accessible. PMID 22900001. doi:10.1371/journal.pone.0041767. 
  11. ^ Alexandra Houssaye, Nathalie Bardet, Jean–Claude Rage, Xabier Pereda Suberbiola, Baâdi Bouya, Mbarek Amaghzaz and Mohamed Amalik (2011). "A review of Pachyvaranus crassispondylus Arambourg, 1952, a pachyostotic marine squamate from the latest Cretaceous phosphates of Morocco and Syria". Geological Magazine. 148 (2): 237–249. doi:10.1017/S0016756810000580. 
  12. ^ Conrad, J.; Rieppel, O.; Grande, L. (2008). "Re-assessment of varanid evolution based on new data from Saniwa ensidens Leidy, 1870 (Squamata, Reptilia)" (PDF). American Museum Novitates. 3630: 1–15. doi:10.1206/596.1. 
  13. ^ Perry, G.; Garland, T., Jr. (2002). "Lizard home ranges revisited: effects of sex, body size, diet, habitat, and phylogeny" (PDF). Ecology. 83 (7): 1870–1885. doi:10.1890/0012-9658(2002)083[1870:LHRREO]2.0.CO;2. 
  14. ^ Clemente, C.J.; Withers, P.C.; Thompson, G.G. (2009). "Metabolic rate and endurance capacity in Australian varanid lizards (Squamata; Varanidae; Varanus)". Biological Journal of the Linnean Society (]). 97 (3): 664–676. doi:10.1111/j.1095-8312.2009.01207.x. 
  15. ^ Greene, Harry W. (1986). Diet and Arboreality in the Emerald Monitor, Varanus prasinus, with Comments on the Study of Adaptation. Chicago: Field Museum of Natural History. OCLC 14915452. Retrieved 12 December 2013. 
  16. ^ Christian, A.; Garland, T., Jr. (1996). "Scaling of limb proportions in monitor lizards (Squamata: Varanidae)" (]). Journal of Herpetology. 30 (2): 219–230. JSTOR 1565513. doi:10.2307/1565513. 
  17. ^ Unidirectional Airflow In The Lungs Of Birds, Crocs And Now Monitor Lizards

External links[edit]

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