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Zhuchengtyrannus Tyrant Of China (TDF Facts)
Zhuchengtyrannus Tyrant Of China (TDF Facts)
Published: 2016/03/25
Channel: The DinoFeed
ASDC - Triceratops vs Zhuchengtyrannus
ASDC - Triceratops vs Zhuchengtyrannus
Published: 2013/02/25
Channel: Allosaurus Rex
PPBA Mapusaurus vs  Zhuchengtyrannus
PPBA Mapusaurus vs Zhuchengtyrannus
Published: 2017/08/16
Channel: Parasaurolophus 67
Zhuchengtyrannus - Video Learning - WizScience.com
Zhuchengtyrannus - Video Learning - WizScience.com
Published: 2015/08/02
Channel: Wiz Science™
Zhuchengtyrannus Magnus
Zhuchengtyrannus Magnus
Published: 2011/10/13
Channel: DINOSTAR5
Yutyrannus chase resounded
Yutyrannus chase resounded
Published: 2016/12/09
Channel: Melvin Spiegelman
Zhuchengtyrannus vs Allosaurus
Zhuchengtyrannus vs Allosaurus
Published: 2013/12/13
Channel: Marc Francois
PPBA Zhuchengtyrannus vs Saurophaganax
PPBA Zhuchengtyrannus vs Saurophaganax
Published: 2015/02/26
Channel: Parasaurolophus 67
accurate theropod dinosaur size comparison 2
accurate theropod dinosaur size comparison 2
Published: 2013/05/15
Channel: TheSPOREREX
Sound effects zhuchengtyrannus
Sound effects zhuchengtyrannus
Published: 2016/03/11
Channel: Cj Magruder
deinonychus123
deinonychus123's Zhuchengtyrannus vs Mapusaurus Resounded
Published: 2014/06/12
Channel: TorvosaurusImperator
PDBA: Yutyrannus Vs Zhuchengtyrannus
PDBA: Yutyrannus Vs Zhuchengtyrannus
Published: 2016/10/08
Channel: Mesozoic StudiosTM
PDW   ALBERTOSAURUS VS DASPLETOSAURUS VS BISTAHIEVERSOR VS GORGOSAURUS VS ZHUCHENGTYRANNUS
PDW ALBERTOSAURUS VS DASPLETOSAURUS VS BISTAHIEVERSOR VS GORGOSAURUS VS ZHUCHENGTYRANNUS
Published: 2015/08/14
Channel: SPINO REX
Zhuchengtyrannus Vs Oxalaia
Zhuchengtyrannus Vs Oxalaia
Published: 2015/09/26
Channel: Behemothic
Zhuchengtyrannus vs Tyrannosaurus
Zhuchengtyrannus vs Tyrannosaurus
Published: 2016/12/30
Channel: SHOTFIN MAKOH
zhuchengtyrannus sounds
zhuchengtyrannus sounds
Published: 2012/03/05
Channel: CAWCarcharo34
Spinosaurus aegypticus vs zhuchengtyrannus magnus (read descr.)
Spinosaurus aegypticus vs zhuchengtyrannus magnus (read descr.)
Published: 2014/12/13
Channel: Giganotosaurus rex pictures
Prehistoric Facts Special #25: Zhuchengtyrannus
Prehistoric Facts Special #25: Zhuchengtyrannus
Published: 2014/06/15
Channel: Christopher Grall
New Carnivore Dinosaur Guide UpDated (Read description for who the dinos are)
New Carnivore Dinosaur Guide UpDated (Read description for who the dinos are)
Published: 2012/04/28
Channel: Allosaurus Rex
Coliseo prehistórico (Deluxe edition): Tarbosaurus vs Zhuchengtyrannus
Coliseo prehistórico (Deluxe edition): Tarbosaurus vs Zhuchengtyrannus
Published: 2016/06/13
Channel: F.P.I.: El Falso Profeta de la Ignorancia
Чжучэнтиранн   Zhuchengtyrannus magnus
Чжучэнтиранн Zhuchengtyrannus magnus
Published: 2016/04/10
Channel: сергей евдокимов
5 Frightening Species of Tyrannosaurus Rex (You
5 Frightening Species of Tyrannosaurus Rex (You've Never Heard Of)
Published: 2017/08/19
Channel: TheTopMan 2.0
UCD paleontologist Dr David Hone on the long lost cousin of T. rex - Zhuchengtyrannus magnus
UCD paleontologist Dr David Hone on the long lost cousin of T. rex - Zhuchengtyrannus magnus
Published: 2011/04/01
Channel: UCD - University College Dublin
2000 views Special:  Brynden Perkins
2000 views Special: Brynden Perkins's Zhuchengtyrannus magnus vs Torvosaurus tanneri Re-Sounded
Published: 2014/08/18
Channel: TorvosaurusImperator
Stunt Rage 2017 Rajasaurus vs Zhuchengtyrannus
Stunt Rage 2017 Rajasaurus vs Zhuchengtyrannus
Published: 2017/07/13
Channel: Gigaston SUPER MAXX!
PBA  Suchomimus vs. Zhuchengtyrannus
PBA Suchomimus vs. Zhuchengtyrannus
Published: 2016/11/15
Channel: Liopleurodon Fragilis
ASDC 3 - Megalosaurus vs Zhuchengtyrannus
ASDC 3 - Megalosaurus vs Zhuchengtyrannus
Published: 2017/04/19
Channel: Liopleurodon Fragilis
Soundscape zhuchengtyrannus vs ankylosaurus
Soundscape zhuchengtyrannus vs ankylosaurus
Published: 2016/07/21
Channel: Cj Magruder
PPBA Tarbosaurus vs Tyrannosaurus
PPBA Tarbosaurus vs Tyrannosaurus
Published: 2015/09/08
Channel: Parasaurolophus 67
ASDC Easter Special - Velociraptor vs Protoceratops vs Oviraptor
ASDC Easter Special - Velociraptor vs Protoceratops vs Oviraptor
Published: 2012/04/07
Channel: Allosaurus Rex
Tyrannosauroid Tribute
Tyrannosauroid Tribute
Published: 2012/04/06
Channel: CKprimeval07
Quick Fights - Tarbosaurus vs Saurophoganax ( 2/3 )
Quick Fights - Tarbosaurus vs Saurophoganax ( 2/3 )
Published: 2017/08/07
Channel: Dinosaur Pictures EntertainmentTM
Paleontology News: All Known Megatheropods
Paleontology News: All Known Megatheropods
Published: 2017/04/28
Channel: The Vividen
nedft 31 Zhuchengtyrannus vs Mapusaurus
nedft 31 Zhuchengtyrannus vs Mapusaurus
Published: 2013/12/30
Channel: deinonychus123
PBA - Zhuchengtyrannus vs. Daspletosaurus vs. Saurophaganax - Entry for SpinoDragon
PBA - Zhuchengtyrannus vs. Daspletosaurus vs. Saurophaganax - Entry for SpinoDragon's Contest
Published: 2016/12/04
Channel: Liopleurodon Fragilis
pckg-Zhuchengtyrannus vs oxalia (round 1 match 16)
pckg-Zhuchengtyrannus vs oxalia (round 1 match 16)
Published: 2013/04/07
Channel: deinonychus123
MDF-Cryolophosaurus vs Zhuchengtyrannus
MDF-Cryolophosaurus vs Zhuchengtyrannus
Published: 2012/07/14
Channel: CeratosaurRevolution
PDFC - Allosaurus vs Camarasaurus
PDFC - Allosaurus vs Camarasaurus
Published: 2013/05/13
Channel: Entersandman1001
Zhunchengtyrannus vs Sinoceratops (final of season)
Zhunchengtyrannus vs Sinoceratops (final of season)
Published: 2017/03/19
Channel: Sebastian Reyna Rivas
OBM - Dinosaur - Zhuchengtyrannus
OBM - Dinosaur - Zhuchengtyrannus
Published: 2016/12/30
Channel: Our Best Moments
PPBA Tyrannosaurus vs Tyrannotitan remake
PPBA Tyrannosaurus vs Tyrannotitan remake
Published: 2017/04/17
Channel: Parasaurolophus 67
zhuchengtyrannus vs therizinosaurs
zhuchengtyrannus vs therizinosaurs
Published: 2012/07/21
Channel: CAWCarcharo34
ASDC 3 - Afrovenator  vs. Zhuchengtyrannus
ASDC 3 - Afrovenator vs. Zhuchengtyrannus
Published: 2017/06/21
Channel: Liopleurodon Fragilis
ASDC - Dilophosaurus vs Postosuchus
ASDC - Dilophosaurus vs Postosuchus
Published: 2013/02/18
Channel: Allosaurus Rex
ASDC - Therizinosaurus vs Tarbosaurus
ASDC - Therizinosaurus vs Tarbosaurus
Published: 2012/04/17
Channel: Allosaurus Rex
PDFC - Carnotaurus vs Triceratops
PDFC - Carnotaurus vs Triceratops
Published: 2017/11/19
Channel: Entersandman1001
Dinosaur News Center: Longest Carnivorous Dinosaur
Dinosaur News Center: Longest Carnivorous Dinosaur
Published: 2010/08/07
Channel: Illiterate Scholar
T.Rex Was Likely an Invasive Species
T.Rex Was Likely an Invasive Species
Published: 2017/06/18
Channel: 21 News
Prehistoric Carnage! (Collab with Paleo Fazbear, MatromX, Etc...Etc... )
Prehistoric Carnage! (Collab with Paleo Fazbear, MatromX, Etc...Etc... )
Published: 2017/01/14
Channel: Chariwade
PPBA Red Eye King vs Vastatosaurus
PPBA Red Eye King vs Vastatosaurus
Published: 2015/02/24
Channel: Parasaurolophus 67
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WIKIPEDIA ARTICLE

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Zhuchengtyrannus
Temporal range: Late Cretaceous, 73.5 Ma
Zhuchengtyrannus mount.jpg
Reconstructed skeleton, with missing bones cast from Tyrannosaurus
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Order: Saurischia
Suborder: Theropoda
Family: Tyrannosauridae
Tribe: Tarbosaurini
Genus: Zhuchengtyrannus
Hone et al., 2011
Type species
Zhuchengtyrannus magnus
Hone et al., 2011
Synonyms

Tyrannosaurus zhuchengensis?
Hu et al., 2001

Zhuchengtyrannus (meaning "Zhucheng tyrant") is an extinct genus of large carnivorous theropod dinosaur known from the Late Cretaceous period of Shandong Province, China. It is a tyrannosaurine tyrannosaurid, and contains a single species, Zhuchengtyrannus magnus.

Discovery[edit]

Diagram showing known remains

Zhuchengtyrannus was first described and named by David W. E. Hone, Kebai Wang, Corwin Sullivan, Xijin Zhao, Shuqing Chen, Dunjin Li, Shuan Ji, Qiang Ji and Xing Xu in 2011 and the type species is Zhuchengtyrannus magnus. The generic name is derived from the word Zhucheng, which refers to the type locality, and tyrant in reference to its phylogenetic position as a tyrannosaurid. The specific name magnus meaning "great" in Latin refers to the relatively large size of Zhuchengtyrannus.[1]

Zhuchengtyrannus is known solely from the holotype ZCDM V0031, a nearly complete right maxilla and associated left dentary (lower jaw, both with teeth) housed at Zhucheng Dinosaur Museum. Casts of the holotype, IVPP FV 1794, are held at the Institute of Vertebrate Paleontology and Paleoanthropology. ZCDM V0031 was collected in situ from the Wangshi Group at Zangjiazhuang quarry, Zhucheng City, dating to the Campanian stage, at least 73.5 million years ago. A second tyrannosaurid dentary (ZCDM V0030) and maxilla (ZCDM V0032) have also been collected at Zangjiazhuang quarry. Even though they were not associated with one another, both specimens are different from other tyrannosaurids, including Zhuchengtyrannus, implying the existence of at least one additional tyrannosaurid from the quarry. Apart from the tyrannosaurid material, specimens of Sinoceratops, hadrosaurids (probably Shantungosaurus) and ankylosaurs were recovered from it.[1] Zhuchengtyrannus was found in an area that was a floodplain in the Cretaceous period and contains one of the highest concentrations of dinosaur bones in the world.[2]

Description[edit]

Estimated size compared to a human.

Zhuchengtyrannus was a large carnivorous theropod, and the holotype has been estimated to have been 10–12 m (33–39 ft) in length and up to 6 short tons (5.4 t) in weight. The holotype dentary is slightly smaller than that of most specimens of Tyrannosaurus rex and slightly bigger than most of Tarbosaurus bataar,[1] but significantly smaller than the corresponding bones of the largest Tyrannosaurus specimen ("Sue").[3]

Zhuchengtyrannus can be distinguished from all other tyrannosaurines by a single autapomorphy, the presence of a horizontal shelf on the lateral surface of the base of the ascending process of the maxilla, and a rounded notch in the anterior margin of the maxillary fenestra. Zhuchengtyrannus also possesses a ventral margin of the antorbital fenestra that lies well above that of the ventral rim of the antorbital fossa. Additionally, the total length of the maxillary fenestra is more than half the distance between the anterior margins of the antorbital fossa and fenestra. Unlike the contemporaneous Tarbosaurus, Zhuchengtyrannus lacks a subcutaneous flange on the posterodorsal part of the jugal ramus of the maxilla, and a ventrally convex palatal shelf that covers the bulges of the roots of the rear teeth in medial view.[1]

Classification[edit]

A bone labeled "Tyrannosaurus Zhuchengensis" on display at the Geological Museum of China.
Illustration from side view.
Teeth labeled "Tyrannosaurus Zhuchengensis" at the Geological Museum of China.
Restored skeletons of Zhuchengtyrannus and Shantungosaurus.

It is possible that several isolated teeth from one of the Zhucheng dinosaur quarries, previously given the name Tyrannosaurus zhuchengensis, belong to this or a related species. The T. zhuchengensis teeth are characterized by serrations that extend all the way to the base of the tooth crown, a feature not seen in any other tyrannosaurine species. All known teeth of Zhuchengtyrannus are too poorly preserved in this area to compare with T. zhuchengensis, but further finds may clarify their relationship.[1]

A phylogenetic analysis published with the description of the tyrannosaurine Lythronax in the journal PLOS ONE recovered Zhuchengtyrannus as the sister taxon of Tarbosaurus. It also suggests that Zhuchengtyrannus and other currently known Asian tyrannosaurids were part of an evolutionary radiation descending from the same North American stem that later gave rise to Tyrannosaurus, recovered as their closest known relative.[4] Below is a cladogram by Loewen et al. in 2013.[4]

Tyrannosauridae


Gorgosaurus libratus



Albertosaurus sarcophagus



Tyrannosaurinae



Daspletosaurus torosus




Two Medicine tyrannosaurid




Teratophoneus curriei




Bistahieversor sealeyi




Lythronax argestes




Tyrannosaurus rex




Tarbosaurus bataar



Zhuchengtyrannus magnus











See also[edit]

References[edit]

  1. ^ a b c d e David W. E. Hone, Kebai Wang, Corwin Sullivan, Xijin Zhao, Shuqing Chen, Dunjin Li, Shuan Ji, Qiang Ji and Xing Xu (2011). "A new, large tyrannosaurine theropod from the Upper Cretaceous of China". Cretaceous Research. 32 (4): 495–503. doi:10.1016/j.cretres.2011.03.005. 
  2. ^ "New dino in same league as T. rex". Sydney Morning Herald. 1 April 2011. Retrieved 3 April 2011. 
  3. ^ Hone D, "So just how big was Zhuchengtyrannus?", archosaurmusings.wordpress.com, 3-4-2011.
  4. ^ a b Loewen, M.A.; Irmis, R.B.; Sertich, J.J.W.; Currie, P. J.; Sampson, S. D. (2013). Evans, David C, ed. "Tyrant Dinosaur Evolution Tracks the Rise and Fall of Late Cretaceous Oceans". PLoS ONE. 8 (11): e79420. doi:10.1371/journal.pone.0079420. PMC 3819173Freely accessible. PMID 24223179. 

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